Today is Betty's birthday, Henry and Betty were supposed to go away for the weekend, however Henry's pregnant girlfriend Charlie shows up in dire need of Henry's attention and ruins their plans. Henry and Betty planned a romantic dinner together, but Charlie is ill and stalls Henry from making it to the restaurant.

Henry leaves a voicemail for Betty, telling her not to go to the restaurant. Betty hears only part of the message and ends up at a table for two waiting for a date that never shows up. Dejected she leaves the restaurant only to run into Gio. Gio takes her on a horse drawn carriage ride through Central Park. They watch fireworks light up the sky. A tempted Betty finally returns home, only to find the house set up for a party and everyone asleep including Henry.

Meanwhile, Alexis and Daniel try to stop their mother Claire from launching her new magazine Hotflash. Alexis financially cuts off her mother so the launch cannot go forward. In retaliation, Claire hires cheap ex-prisoners to staff the start-up.

Rene, Whilhelmina's sister, reveals that she is afraid of candles. People begin to become suspicious of her erratic behavior.

Wilhelmina is trying to break up Rene and Daniel, she told Daniel to ask her about Stonybrook, when he did she just replied it is a school I went to. Rene warned Wilhelmina to stay out of her business because she is not the only one with secrets, meanwhile Christina is eavesdropping and hears it all jotting down the word Stonybrook. Later Rene and Marc share a chocotini and Marc spills his guts about Wilhelmina's stealing Brandford's sperm and having Christina impregnated. Rene confronts Wilhelmina with what she found out, she tells Wilhelmina to back off Daniel or she will tell her everything she knows.

Rene ends up telling Daniel she had a breakdown at Stonybrook and now takes antidepressants, Daniel said it doesn't make him love her any less and asks her to move in. Meanwhile Wilhelmina had Marc substitute her antidepressants with fat blocker pills.

Betty has been working non stop all week helping Daniel prepare for the big Christian Sirano meeting, Rene is driving her nuts with incessant calls and text messages. Betty later catches her changing Daniel's schedule around, Daniel walks in and sees them fighting over the appointment book, Rene then accuses Betty of being in love him, now Daniel is paranoid and reading things into every move Betty makes.

Christian Sirano makes his grand entrance with his entourage of beautiful models, when Rene barges in, making a scene, wanting to know if they are Daniels whores!

Meanwhile Wilhemina has spoken with Rene's therapist to see what symptoms she might exhibit if she were experiencing another break down, later she schedules a lunch with Rene, however, it happens to be at a restaurant that holds bad memories for Rene, who freaks out and leaves when the waiter shows up with a cake full of candles.

Daniel and Wilhelmina then meet with her doctor again who describes her as suffering from borderline personality disorder, he assures them that Rene would not intentionally hurt Daniel, but if Rene thinks another woman is involved then that woman would be the one in danger.

Christina and Betty do some snooping at Wilhemina's, they stumble on a file containing Rene's psychiatric report which details the fiery murder of her last boyfriend. Betty rushes to Daniel's apartment, only to be greeted by Rene in a room full of candles. The two struggle and the curtains catch fire, Betty manages to talk Rene down before Daniel and Wilhelmina arrive.

Henry tells Betty about Charlie's panic attack at the airport, the doctor does not want her to travel until after the baby is born. Henry later confronts Gio telling him to stay away from Betty, Henry takes a wild swing and catches Gio in the nose.

Henry and Betty were enjoying a romantic dinner when Charlie barged in and started eating their dinner, Charlie ruined their lunch plans the next day as well, she claimed her yoga partner couldn't make it so Henry was going to go, Betty decided she would go instead much to Charlie's displeasure, they end up getting kicked out. Betty then plans a baby shower for her, Charlie is quite rude and off putting during the shower. Betty ends up telling her off & Charlie's water breaks in the middle of the argument. Betty and Charlie have an honest talk while waiting for the ambulance, Charlie admits she is afraid of raising a child alone, by the time the ambulance arrives it's too late to bring her to the hospital, she will deliver her baby at Betty's.

Christina and Betty enjoy a lunch in the park together, Christina finds out she is carrying the late Mr Meads child, upset she runs to Stuart, who calls Wilhelmina and asks for more money.

Everyone is concerned about Daniel since his break up with Rene, Betty scheduled him an appointment with a therapist, Daniel hits on the therapist and gets kicked out. He returns later and starts to honestly talk about Rene, however they end up sleeping together, which prompts his therapist to call her therapist.

Wilhelmina plans a special announcement, however it is put on hold when Christina ends up missing. Christina shows up at Wilhelmina's office and makes her watch the ultrasound tape while promising to love this child, she takes that ultrasound and uses it to announce to Daniel and his mother that she is having the elder Meads child!

Hilda goes to the gym teacher about Justin not participating in gym class, he is not what she expected, he is hot! She offers to cut his hair, but before she begins she cuts his ear, sending him running from the shop. Hilda goes to see the coach to apologize and while there he offers to let Justin help out with the cheerleaders and he will pass him.

Betty is trying to get over Henry, since he, Charlie and the baby went back to Tuscon. Meanwhile Gio tells Betty to admit that she likes him, he grabs her and kisses her, she then slaps him, however the kiss was caught on tape and sent to everyone in the office.

Wilhelmina is back at Mode, her first plan is to divide the family, she sits in on a meeting and sets her plan in motion by pitching an idea to Alexis while Daniel was in the men's room. Daniel tries to do damage control by going on TV, however someone calls the show and discredits him. Meanwhile Alexis reinstates Wilhelmina as creative director of Mode magazine, despite protests from both Daniel and her mother.

Amanda gets ready for her reality show with her father, Gene Simmons. During the show Amanda notices that Gene's Tweety bird tattoo is fake, giving rise to the question, are you my father? Amanda is devastated when she finds out that he was just using her for a publicity stunt.

Hilda flirts with Tony the hot gym teacher, she volunteers to chaperone for the school dance, however when she gets sick Betty goes in her place where she runs into Gio again. Later in the evening Justin approaches the coach and tells him how much his mom likes him. Tony later stops by with some soup for Hilda and invites her on a date when she is feeling better.

Daniel has been putting in a lot of hours at Mode, meanwhile a young French boy visits Daniel and with the help of a translator he tells him he is his father. Daniel couldn't handle it and left to go party with some models.

Wilhelmina steps in a takes over during a big meeting, later she approaches Alexis to suggest she do what is best for Mode and find Daniel another position in the company.

Gio tries to teach Betty how to play softball for the Mode softball team, after much practice Betty finally gets it. Gio invites Betty to go to Rome with him, Betty says yes, meanwhile Henry shows up to ask Betty to marry him and move to Tuscon. Daniel presents Betty with two first class tickets to Rome, Betty breaks down and tells him about Henry's proposal, Daniel suggests she go to Rome.

Alexis asks Daniel to step down and take a leave of absence due to the bad press surrounding Daniel, concerning the story about abandoning his son.

The coach tells Hilda he is not exactly single, breaking her heart, later she decides to take a chance on him.

It's time for the big softball game between Mode and Elle magazine with big names like Naomi Campbell, Gio and Henry meet at the game and both want Betty's answer, while Wilhelmina announces to Daniel she is now editor and chief.

Betty gets knocked out at the game, while she is unconscious she sees the two directions her life could go in depending on which man she says yes to, when she comes to she decides what she is going to do.