It all started with the little anime site that could, 4-chan. It's a forum or messaging board that focuses on images. Each folder has its own topic for messages/images, be it anime, cute anime, cosplay, mecha, etc.

You're probably thinking specifically of /b/, which is the forum for random images. /b/ is the virtual gutter of the internet, where pictures are tossed about for maximum shock value. /b/ is the purveyor of the vilest of the vile images lurking on the internet, such as tubgirl, goatse, and 2 girls 1 cup. It’s like a rotten.com for the 21st century. Conversely, it is the source for nearly every popular meme on the internet current and former, from Rickrolling to LOLcats to All Your Base.

Some /b/-tards, as they are called, operate as Anonymous, a loose group of people who share similar disdain towards certain groups of people, such as people who take the Internet way too seriously, pro-censorship politicians, the RIAA/MPAA, and the Church of Scientology. Anonymous will occasionally prank or torment these groups in order to inconvenience them or make them furious so they can sit back and laugh at how angry they got.
Source(s):
http://en.wikipedia.org/wiki/4chan
http://extremelylame.com/what-is-anonymous/


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