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Twinkies are baked for 10 minutes, then cream is injected through three holes in the browned top of the cake, which is flipped during packaging so the rounded part becomes the top.
FYI Hostess makes more than 500 million Twinkies every year. That's 1,000 every minute. They also use 40,000 miles of cellophane to wrap all those Twinkies. Yum!
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According to this source:
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Three injectors simultaneously pierce the bottom of the cake, filling it with three "globs" of filling. If you look at the bottom of a Twinkie, you can usually see where the filling was injected.
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